| Maynard Smith, J. 1965. The evolution of alarm calls. The American Naturalist, 99:59-63. |
....are rare, and that greater pursuit costs to the predator will allow cheaper signals by the prey. 1 Introduction Alarm calling behavior has provided a persistent puzzle to behavioral biologists. Why should an individual risk its own survival to produce costly warnings of a predator s presence (Maynard Smith, 1965) Evolutionary biologists have typically invoked kin selection and or reciprocal altruism to explain such behaviors (reviewed by Hauser 1996, pp. 413 433) While a calling individual may su er some direct cost, it may also reap inclusive tness bene ts by warning relatives of danger, or anticipate ....
Maynard Smith, J. 1965. The evolution of alarm calls. The American Naturalist, 99:59-63.
....spatial organization Another possible solution involves changing the locus at which selection is presumed to operate. Ethologists have argued that this locus, rather than being at the level of the individual, is better understood as being at the level of the gene (Hamilton, 1963; Hamilton, 1964; Maynard Smith, 1965; Dawkins, 1976) Because a given gene will exist in more that one individual, an apparently altruistic act may well be selfish with respect to a gene. In particular, related individuals will generally expect to share genes in common (50 for siblings and parent child relationships, reducing by ....
....transmitter. The standard deviation of the spatial distribution is 5 locations in the population. Four meanings and four signals are used. Results are averaged over ten simulation runs. existence of alarm calls, given that the degree of relatedness is sufficient (Hamilton, 1963; Hamilton, 1964; Maynard Smith, 1965). This theoretical work is supported by field evidence in animals such as the Belding s ground squirrel, where alarm call frequency is related to the expected presence of related individuals (Sherman, 1977) Following the structure used in the spatialized version of the prisoner s dilemma, similar ....
Maynard Smith, J. (1965). The evolution of alarm calls. American Naturalist 99, 59--63.
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J. Maynard-Smith. The evolution of alarm calls. American Naturalist, 99, 1965.
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